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Are you referring to the screen at the top of the center console? If so, there are 2 different mpg numbers that can be displayed.
One of them is the mpg for the tank of gas since last fill up. This display will have the letters AVG (for average) before the number. It is supposed to reset automatically when you fill the gas tank all the way up. There is no manual reset that I know of.
The other mpg reading is the current fuel consumption at that moment. There is no reset on this at all, it is a constantly changing number.
Toyota and Lexus Join Mille Miglia For The First Time
Slideshow: A five-car lineup spanning more than five decades of Toyota performance and engineering will tackle one of Italy's most celebrated automotive routes.